Layer remains when slide restarts

Hi. I'm having an issue with a slide with multiple layers and a custom replay button using a client's requirements/template. Initially the buttons for the layers are disabled but change to normal when the replay button appears (at a cue point at the end of the timeline). At that point the user can click on any of the buttons (boxes) and appropriate layer appears. When the replay button is clicked, the slide restarts but the last layer clicked remains displayed on screen (the screen should be blank). I've tried several things, but I can't find the setting/trigger/state/variable that is causing this behavior. Help/insight would be appreciated. 

Story file attached. Review link: https://360.articulate.com/review/content/2c34d117-79da-4b53-9747-3f420d5a0d37/review